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An alien invasion is set to happen in the city of Akiba. The town's only hope is the Papillion Soldiers, who now lead mundane lives and do not remember their past heroics.
Papillion Rose tries to convince Anne that she is Lily and that Shizuka is Margarette. Meanwhile, the Dark Advisor is sent down to Earth to speed up the Forgotten Ones' mods.
The Dark Advisor declares war on Earth by mobilizing ground units against the Papillion Soldiers with hopes that the people of Earth will surrender.
The invaders send out a three-headed dragon. Later, the girls visit Shinjuku to relax, learning about Papillion Dahlia.
The Papillion Soldiers realize that Dahlia wields the legendary Kusanagi and is under the enemies control. To get Dahlia back on their side, they must fight her for the sword.
In their efforts to save Rose from eternal doom, the girls learn a key function of the Kusanagi's power.