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The story of the Rozen Maidens is retold, giving rise to a new tale of the eternal power struggle that is the Alice Game.
In an alternate reality, Jun decides to "Not Wind It", and continues to live a normal, if somewhat sad life. Things change when he gets a strange package in the mail.
Jun continues to receive messages from his younger self, urging him to complete "Shinku" at all costs.
Jun wakes up to find a completed Shinku. With a few twists, he brings her to life and embroils himself further in the Alice Game.
Suigintou has made her way to Shinku, and the melancholic journey she underwent to get there is revealed.
To get back to the Jun who wound, Shinku and Suigintou call a truce. Elsewhere, adult Jun is humiliated.
Hoping to chance his world, adult Jun begins working on a "new doll" behind Shinku's back. Elsewhere, young Jun is at a loss with what to do next.
Adult Jun's new doll is only complete, missing only its head. Later, the play he is helping on runs into some trouble.
Kirakisho shows her true self, and Adult Jun, Suigintou, and Shinku have no choice but to fight for their lives.
A familiar face returns, and with some help, attempts to reclaim what is hers. Young Jun and Adult Jun finally meet.
As Souseiseki sets out to restart the big clock, she strikes a deal with Suigintou, one that she keeps from her twin. Young Jun finally finds the true Shinku.
With Kirakisho seemingly defeated, the Rozen Maidens and Juns part ways, but not before Suigintou sets off to look for Megu.
Just who didn't wind the key returns to his normal life…or so he thinks. His story isn't over, however, and Shinku and the other dolls approach this new challenge together.